Coldfall Wood
Report on a ground invertebrate survey, 30 April 2009 to 29 April 2010
Edward Milner B Sc, MIEEM
5. List of other invertebrates recorded from Coldfall Wood
1. Introduction
Coldfall Wood (14ha), one of the very few fragments of ancient woodland left in north London. It was originally part of the extensive area of ancient woodland north of London that existed in medieval times. This covered much of what is now the built-up areas of Finchley, Muswell Hill and parts of Barnet, and Coldfall Wood, along with Big Wood, and parts of Coppetts Wood are some of the small fragments that are left.
Until the 1940’s Coldfall was managed as a (hornbeam) coppice wood with oak standards, as were many other woods around London, managed to provide fuel for the metropolis. Colfall Wood is predominantly hornbeam with a little hazel, some wild service, holly, birch, rowan and occasional crabapple. There are wild cherry suckers but no mature cherry trees. Most of the wood was probably cut primarily for firewood but some or all of it may have been converted to charcoal on site. Coppicing was discontinued after the Second World War and the hornbeams have grown up so that by the 1990s over most of the wood the canopy had closed, allowing very little light to reach the woodland floor after early spring. As a result the ground flora, and associated invertebrates were substantially restricted.
There is a level area in the south-western part of the wood which could have been the charcoal-burners base. It was a clearing; this can be inferred from the number of birch trees which replace hornbeam in the immediate vicinity. Birches are a pioneer species which do not form a dense canopy, and the trees being very much younger than most of the other trees in the wood. Today this area has grass with bracken because the tree canopy is thinner.
In recent years coppicing of the hornbeam (and some wild service) has been reinstated on a piecemeal basis. Three areas were cut in the early 1990’s, and finally a large swathe down most of the length of the main stream was cut in the winter of 2006-7. Some pitfall-trapping was done in the early 1990’s and a preliminary species list of 104 spp. resulted. This included (from a damp area in one of the re-copppice cuts) a rare money spider (Porrhomma cambridgei) which was named as a species new to science on the basis of specimens from Coldfall Wood and five other UK sites.
As a result of the re-coppicing some plants not previously recorded from the wood have appeared as young seedlings, and the vegetation structure of these coppiced areas is very different from other parts of the wood. Studies by David Bevan (formerly the Conservation Officer for Haringey) have shown that since the most recent coppicing the number of new plant species has increased enormously, rising from 55 spp. before this cut to 230spp. by 2009. Many of these will have arrived on the wind or been brought in by birds, and some are aliens and garden escapes. However it is clear that quite a number have arisen from seeds buried in the seedbank, and of these some are rare species only known from one or two other sites in the whole of Middlesex.
Invertebrates may take longer to re-establish themselves, especially plant-feeders which need to have well-established food-plants. or appear because they are associated with some of the plants, usually as plant-feeders. Spiders are more dependant upon the complexity of the plant structure rather than the particular plant species (being carnivorous rather than plant-feeding), but the newly coppiced areas, especially the damper areas and streamside vegetation will gradually mature providing a complexity of structure that will offers a wide range of niches for different spider species.
2. The Survey
This survey concentrated on the newly coppiced area (5 trap sites) with some control sites elsewhere in the wood including the grassy Lovers Lawn. A list of the trap sites is given in Table 1 (see also Map).
Table 1: Habitat Compartments and trap-sites
| Compartment | Trap-site | Habitat | Ground vegetation | No of spider spp. recorded |
E | 01 | Damp, above top pool, mainly Juncus and part shaded by Leycesteria formosa | Dense | 14 |
| E | 02 | Dry, part shaded, edge of new coppice under Carpinus | Sparse | 14 |
| D | 03 | Dry, in ground vegetetation (Hedera helix) at edge of new coppice | Dense | 14 |
| D | 04 | Dry, bare ground (1 m from stream) with Juncus in new coppice area | Sparse | 25 |
| D | 05 | Damp, edge of dense Rubus and tree seedlings area in largest new coppice area | Dense | 22 |
| H | 06 | (Control) Leaf litter under dense Carpinus canopy | Absent | 13 |
| H | 07 | Dry, grassy area with Pteridium; old charcoal-burners clearing | Sparse | 25 |
| H | 08 | Dry, edge of Rubus patch at base of large oak, in early (1991-2) coppice cut | Sparse | 25 |
| H | 09 | Among dense ground vegetation (Hedera) in dense shade of hornbeam coppice from early cut (1990-91) | Dense | 12 |
| J | 10 | Dry, grassy area (Lovers Lawn) | Dense (grass) | 27 |
Pitfall traps were set in standard trios with a small quantity of ethylene glycol (holding agent) and commercial detergent (wetting agent). These compounds may act as an attractant for some species and discourage others but this standard form has been used throughout London in previous surveys and presumably the same bias (if any) occurs each time. The traps were emptied and the catch sorted on a monthly basis.
The survey began when the traps were first set on April 30 2009 and will continue until April 29 2010. This interim report covers the period April 30 2009 to December 31st 2009, but updated lists of species will be provided at the conclusion of the 12-month survey period.
Spiders were identified by the author, beetles by Mr N Heal, the pseudoscorpions by Dr G Legg, and bugs (Hemiptera) by Mr T Bantock.
3. Results and interpretation
In total 56 spp. of spiders, 79 spp. of beetle, and 3 species of pseudoscorpion have been recorded during the study period. The totals for these groups for the wood as a whole are: spiders 119spp, beetles 106 spp. (including three Nationally Notable Species) and pseudoscorpions 3 spp. Complete lists of all invertebrate records for the wood are appended.
Before the present survey while few beetles had been recorded from the wood, spiders were already better known (104 spp. recorded). This was an incomplete list produced from some limited collecting and sporadic pitfall-trapping by the author. The present survey has so far produced 15 spider spp. not previously recorded for the wood.
The main conclusion to be drawn is that as a result of the new coppicing the wood provides a richer and more complex habitat for invertebrates, and this process will probably continue for many years. It is difficult to estimate what the species richness of spiders or beetles will be like in ten or fifteen years from now, but it is to be expected that both would be considerably greater than today’s figures indicate.
(a) Spiders
Most of the spiders in the wood are money spiders (Linyphiidae) which made up 64% of the catch during the survey. This was as expected. The part of the habitat sampled is the ground level, including the leaf-litter zone and under shaded or part-shaded conditions money spiders are the most abundant and diverse spiders in London woodlands, where heavy shade predominates in neglected hornbeam coppice.
The spider data given in Table 1 shows that the trap-sites divide clearly into two groups; the poorer sites with 12-14 spp. ( 01,02,03, 06 and 09) and the richer sites with 22-27 spp. (04,05,07,08,10). The mean number of species for the second group is double the mean number for the first group. In other words the second group (mostly less shaded) have a very much richer spider fauna than the first group (more heavily shaded). This may change in future as the habitats mature.
The two damper sites (02 and 05) are both somewhat shaded and have so far not produced any species characteristic of damp habitats, but such species can be expected to appear over time. The open areas of the new cut (02 and 03 especially) may also produce more non-woodland spiders taking advantage of the ‘woodland edge’ habitat.
It was hoped that the spider survey might show that new spiders had arrived in the recently cut coppice zone in a similar way to the plants. However this may take some time several years at least, and much longer than the present survey. Details about the new species recorded are given in the next section. Perhaps if trapping is to continue, one or two more trap-sites in open parts of the new cut should be considered.
The newly recorded spider species
A few of these spiders may be associated with the new coppice cut; these are the seven species marked *. Others are recorded for the first time and reflect the fact that the wood has not been much investigated for spiders apart from the pitfall-trapping.
Harpactea hombergi (Dysderidae)(six-eyed spiders)
This is not a ground-dwelling species but lives under the bark of trees and may move between trees sometimes. Recorded from site 02.
Oonops pulcher (Oonopidae)(tiny six-eyed spiders)
This tiny orange spider has a similar habit to H hombergi and is not normally found in pitfalls. Recorded from site 02.
*Walckenaeria atrotibialis (Linyphiidae)(money spiders)
This small money spider is a grassland/low vegetation species, not a woodland species. Recorded from sites 05, 07
Dismodicus bifrons (Linyphiidae)
Normally found in damp places outside woodland. Recorded from site 10.
*Oedothorax retusus (Linyphiidae)
A pioneer of damp grassland and low vegetation; not a woodland species. Recorded from sites 04 and 05.
Erigonella hiemalis (Linyphiidae)
A tiny woodland species that is often more active in the winter months. Recorded from sites 05 and 07.
*Meioneta saxatilis (Linyphiidae)
A pioneer of various habitats. Recorded from site 10.
Tapinopa longidens (Linyphiidae)
A woodland spider; not very common in London. Recorded from site 10.
Lepthyphantes mengei (Linyphiidae)
A small woodland species. Recorded from sites 02, 04, 07, and 09.
*Pardosa prativaga (Lycosidae)(wolf spiders)
A spider of open ground, not a woodland species, a pioneer. Recorded from sites 01, 04, 05.
*Pisaura mirabilis (Pisauridae)
A spider of tall vegetation and dense grass; not a woodland species. Recorded from site 08.
Clubiona comta (Clubionidae)(mouse spiders)
A woodland species that lives much of the summer in the canopy and which migrates up the trees in spring and down in winter although many remain in small silk cells under bark. Recorded from sites 03 and 04.
*Tibellus oblongus (Philiodromidae) (long-legged crab spiders)
A straw-coloured spider of long grass and open habitats; not a woodland species. Recorded from site 10.
Ozyptila praticola (Thomisidae) (crab spiders)
A spider of leaf-litter in dense woodland. Recorded from sites 02 and 08.
*Ozyptila simplex (Thomisidae)
A spider of open ground; not a woodland species. Recorded from site 08.
(b) Beetles
The most interesting beetle recorded during the survey is the very rare Lesser Glowworm, Phosphaenus hemipterus (Lampyridae). As a recent paper in the British Wildlife magazine (De Cock et al., 2009) has shown, while there is some doubt as whether it is a genuine native species, there have been very few records for the whole country. Since the first record in 1868 they detail just 19 records, all in the south east of England (and none north of the Thames). Apparently a specimen was also found in 2009 at Orpington in south London, but we have had one adult and three larvae from Coldfall Wood, at least some of these from site 05 in the middle of the newly coppiced area.
Three other beetles found are Nationally Notable B species: two carnivorous species: a ground beetle Platyderus depressus, a rove beetle Anotylus mutator; and a plant-feeding click-beetle Athous campyloides, associated with grasses as food-plants.
Most of the other beetles species found in this survey are widespread and common in London. Since most beetles are extremely mobile their occurrence in pitfall traps is less easy to interpret than spider results. However, plant-feeding species are usually found near their food-plants. So far, among the small number of plant-feeding beetles recorded, most are general feeders being found on a wide range of plant hosts. However one tiny weevil, Aracharius salicivorus a willow-gall beetle, has been trapped. This species is restricted to goat willow, one of the plants that has appeared in the wood for the first time in the newly coppiced zone.
In future seasons many further species can be expected to appear as the habitat matures and plant-feeding species follow the new plants which have arrived. This will be especially the case with those beetles which are restricted to certain plant species by their feeding preferences – many plant genera have specific weevils and flower-beetles which are restricted to them. So far, the lists for these families (Chrysomelidae, Apionidae and Curculionidae) are very poor (totalling only 13 spp.) and over the next few years the diversity of these beetles can be expected to increase to at least 50 and possibly 100.
4. Reference
De Cock, R, Horne, J, and Tyler, J. 2009. The Lesser Glow-worm in Britain: native or newcomer. British Wildlife 21 (1): 39-43.
5. Species lists
Key for all following lists:
First column shows National Conservation Status
RDBK = high conservation status (rare) but precise status unknown
Na and Nb = Nationally Notable A & B
Loc. = local in distribution
Comm. = common.
Date given is that of the first record
Serial is the author’s reference no. for the record
Last column is initials of author
1. List of spiders recorded during the present survey
DYSDERIDAE (Six-Eyed Spiders)
Comm Harpactea hombergi 30/05/2009 016759/006
OONOPIDAE (Six-Eyed Spiders)
Comm Oonops pulcher 30/06/2009 016791/012 em
THERIDIIDAE (Tangle Web Spiders)
Comm Theridion mystaceum 31/07/2009 016836/013
Loc Enoplognatha thoracica 30/05/2009 016754/005 em
Comm Robertus lividus 30/05/2009 016762/007
LINYPHIIDAE (Money spiders)
Comm Ceratinella brevipes 30/05/2009 016762/011
Comm Walckenaeria acuminata 30/05/2009 016754/002 em
Loc Walckenaeria atrotibialis 30/06/2009 016790/012 em
Loc Dicymbium brevisetosum 31/07/2009 016836/009
Comm Gongylidium rufipes 30/05/2009 016758/017
Loc Dismodicus bifrons 30/05/2009 016762/010
Comm Maso sundevalli 30/05/2009 016754/010 em
Comm Oedothorax fuscus 30/11/2009 016954/004
Comm Oedothorax retusus 30/05/2009 016761/009
Comm Tiso vagans 30/05/2009 016754/004 em
Comm Monocephalus fuscipes 30/05/2009 016755/003 em
Comm Gongylidiellum vivum 30/05/2009 016762/006
Comm Micrargus herbigradus s s 30/05/2009 016754/011 em
Comm Erigonella hiemalis 30/05/2009 016757/012
Comm Diplocephalus cristatus 28/08/2009 016853/003
Comm Diplocephalus latifrons 30/05/2009 016754/007 em
Comm Diplocephalus picinus 30/05/2009 016754/006 em
Comm Erigone dentipalpis 30/05/2009 016754/008 em
Comm Erigone atra 30/05/2009 016758/014
Comm Meioneta saxatilis 30/06/2009 016800/006 em
Comm Microneta viaria 30/05/2009 016754/003 em
Comm Centromerus sylvaticus 30/12/2009 016983/003
Comm Centromerita bicolor 30/11/2009 016958/005
Comm Saaristoa abnormis 30/05/2009 016762/012
Comm Macrargus rufus 30/05/2009 016759/001
Comm Bathyphantes gracilis 30/05/2009 016760/006
Comm Diplostyla concolor 30/05/2009 016755/002 em
Comm Drapetisca socialis 30/10/2009 016923/006
Comm Tapinopa longidens 30/10/2009 016920/003
Comm Lepthyphantes minutus 30/10/2009 016922/006
Comm Lepthyphantes tenuis 30/05/2009 016754/009 em
Comm Lepthyphantes zimmermanni 30/05/2009 016756/009 em
Comm Lepthyphantes mengei 30/05/2009 016757/010
Comm Lepthyphantes flavipes 30/05/2009 016754/012 em
Loc Lepthyphantes pallidus 30/05/2009 016757/014
Comm Linyphia hortensis 30/06/2009 016792/004 em
TETRAGNATHIDAE (Orb Web spiders)
Comm Pachygnatha degeeri 30/05/2009 016754/001 em
Comm Metellina segmentata 30/10/2009 016922/008
LYCOSIDAE (Wolf Spiders)
Comm Pardosa pullata 30/05/2009 016758/008
Comm Pardosa prativaga 30/05/2009 016758/006
Comm Pardosa amentata 30/05/2009 016758/005
Comm Pardosa saltans 30/05/2009 016758/004
Comm Alopecosa pulverulenta 30/05/2009 016758/003
Comm Trochosa terricola 30/05/2009 016756/003 em
PISAURIDAE (Nursery Web Spiders)
Comm (imm) Pisaura mirabilis 30/10/2009 016923/007
CLUBIONIDAE (Mouse Spiders)
Comm Clubiona terrestris 30/05/2009 016756/004 em
Comm Clubiona comta 30/05/2009 016756/005 em
ZORIDAE
Comm Zora spinimana 31/07/2009 016828/007
PHILODROMIDAE (Elongate Crab Spiders)
Comm Tibellus oblongus 30/05/2009 016762/013
THOMISIDAE (Crab Spiders)
Comm Ozyptila praticola 31/07/2009 016836/011
Loc Ozyptila simplex 30/06/2009 016791/006 em
2. List of all spiders recorded from Coldfall Wood
DYSDERIDAE (Six-Eyed Spiders)
Comm Harpactea hombergi 30/05/2009 016759/006
OONOPIDAE (Six-Eyed Spiders)
Comm Oonops pulcher 30/06/2009 016791/012 em
NESTICIDAE
Loc Nesticus cellulanus 09/07/1994 004186/009 em
THERIDIIDAE (Tangle Web Spiders)
Comm Anelosimus vittatus 06/06/1993 003837/003 em
Loc Achaearanea lunata 15/05/1991 003310/001 em
Comm Theridion sisyphium 04/06/1991 003365/001 em
Comm Theridion varians 04/06/1991 003365/004 em
Comm Theridion mystaceum 18/05/2008 016320/027
Comm Neottiura bimaculatum 18/05/2008 016320/021
Comm Paidiscura pallens 17/07/1993 003804/009 em
Comm Enoplognatha ovata s s 28/05/1990 002927/004 em
Loc Enoplognatha thoracica 13/05/1991 003324/012 em
Comm Robertus lividus 16/07/1993 003849/007 em
LINYPHIIDAE (Money spiders)
Comm Ceratinella brevipes 04/06/1991 003358/012 em
Comm Walckenaeria acuminata 02/02/1991 003202/003 em
Comm Walckenaeria antica 02/05/1994 004120/012 em
Comm Walckenaeria cucullata 31/03/1991 003331/005 em
Loc Walckenaeria atrotibialis 30/06/2009 016790/012 em
Comm Walckenaeria nudipalpis 28/02/1993 003722/007 em
Loc Dicymbium brevisetosum 22/11/1992 003633/009 em
Comm Entelecara acuminata 13/07/1992 003617/011 em
Loc Entelecara erythropus 28/05/1990 002927/008 em
Comm Gongylidium rufipes 01/06/1986 000664/002 em
Loc Dismodicus bifrons 30/05/2009 016762/010
Comm Hypomma bituberculatum 18/05/2008 016320/032
Comm Maso sundevalli 04/06/1991 003365/002 em
Comm Pocadicnemis pumila 09/07/1994 004186/010 em
Comm Pocadicnemis juncea 21/07/1991 003398/009 em
Comm Oedothorax fuscus 12/08/1991 003396/011 em
Comm Oedothorax retusus 30/05/2009 016761/009
Loc Cnephalocotes obscurus 04/06/1991 003358/013 em
Comm Tiso vagans 02/12/1984 000268/005 em
Comm Monocephalus fuscipes 02/12/1984 000268/004 em
Comm Gongylidiellum vivum 19/12/1992 003638/005 em
Comm Micrargus herbigradus 06/01/1991 003227/006 em
Comm Erigonella hiemalis 30/05/2009 016757/012
Comm Diplocephalus cristatus 12/08/1991 003396/013 em
Comm Diplocephalus permixtus 02/12/1984 000268/006 em
Comm Diplocephalus latifrons 02/12/1984 000268/001 em
Comm Diplocephalus picinus 28/05/1990 002927/001 em
Comm Araeoncus humilis 27/03/1993 003735/011 em
Loc Panamomops sulcifrons 06/06/1993 003835/008 em
Loc Lessertia dentichelis 01/01/1993 003669/006 em
Loc Milleriana inerrans 12/08/1991 003392/009 em
Comm Erigone dentipalpis 13/04/1985 000333/004 em
Comm Erigone atra 02/02/1991 003202/005 em
RDBK Porrhomma cambridgei 27/03/1993 003731/012 em
Loc Agyneta decora 12/08/1991 003396/016 em
Comm Meioneta rurestris 02/12/1984 000268/002 em
Comm Meioneta saxatilis 30/06/2009 016800/006 em
Comm Microneta viaria 02/12/1984 000268/003 em
Comm Meioneta innotabilis 18/05/2008 016320/033 em
Comm Centromerus sylvaticus 06/01/1991 003227/005 em
Comm Centromerus dilutus 10/02/1985 000299/005 em
Comm Centromerita bicolor 06/01/1991 003227/004 em
Comm Saaristoa abnormis 12/08/1991 003396/015 em
Loc Saaristoa firma 28/10/1992 003626/004 em
Comm Macrargus rufus 10/02/1985 000299/001 em
Comm Bathyphantes gracilis 13/04/1985 000333/007 em
Comm Bathyphantes parvulus 21/07/1991 003394/010 em
Comm Diplostyla concolor 10/02/1985 000299/004 em
Comm Drapetisca socialis 02/10/1991 003432/003 em
Comm Tapinopa longidens 30/10/2009 016920/003
Comm Labulla thoracica 21/10/1992 003549/004 em
Comm Stemonyphantes lineatus 30/11/1991 003420/008 em
Comm Lepthyphantes minutus 02/12/1984 000268/007 em
Comm Lepthyphantes tenuis 13/04/1985 000333/005 em
Comm Lepthyphantes zimmermanni 13/04/1985 000333/006 em
Comm Lepthyphantes mengei 30/05/2009 016757/010
Comm Lepthyphantes flavipes 25/05/1985 000353/005 em
Comm Lepthyphantes ericaeus 12/08/1991 003396/008 em
Loc Lepthyphantes pallidus 25/05/1985 000353/006 em
Comm Helophora insignis 30/11/1992 003653/001 db
Comm Linyphia triangularis 18/05/2008 016320/004
Comm Linyphia hortensis 06/06/1993 003837/005 em
Comm Neriene montana 13/05/1991 003322/007 em
Comm Neriene clathrata 27/03/1993 003735/007 em
Comm Neriene peltata 04/06/1991 003365/003 em
TETRAGNATHIDAE (Orb Web spiders)
Nb Tetragnatha pinicola 06/06/1993 003833/003 em
Loc Tetragnatha montana 06/06/1993 003837/006 em
Comm Pachygnatha clercki 18/05/1993 003807/001 em
Comm Pachygnatha degeeri 06/01/1991 003227/001 em
Comm Metellina segmentata 28/06/1992 003463/001 em
Comm Metellina mengei 06/06/1993 003837/004 em
ARANEIDAE (Orb Web spiders)
Comm Araneus diadematus 25/09/1993 003850/001 em
Loc Araniella opisthographa 18/05/2008 016320/025
Loc Cyclosa conica 02/05/1994 003990/001 em
LYCOSIDAE (Wolf Spiders)
Comm Pardosa pullata 25/04/1991 003341/009 em
Comm Pardosa prativaga 30/05/2009 016758/006
Comm Pardosa amentata 01/06/1986 000664/005 em
Comm Pardosa saltans 13/05/1991 003324/008 em
Comm Alopecosa pulverulenta 25/04/1991 003341/002 em
Comm Trochosa ruricola 26/09/1992 003465/002 em
Comm Trochosa terricola 31/03/1991 003342/001 em
PISAURIDAE (Nursery Web Spiders)
Comm (imm) Pisaura mirabilis 30/10/2009 016923/007
AGELENIDAE (House Spiders & allies)
Loc Tegenaria gigantea 26/09/1992 003467/006 em
Loc Tegenaria agrestis 21/10/1992 003549/003 em
HAHNIIDAE
Loc Hahnia helveola 06/01/1991 003226/002 em
DICTYNIDAE (Small Blue Web Spiders)
Comm Dictyna uncinata 06/06/1993 003841/005 em
AMAUROBIIDAE (Blue Web Spiders)
Comm Amaurobius ferox 18/05/2008 016320/009
LIOCRANIDAE
Comm Phrurolithus festivus 29/06/1991 003397/006 em
Loc Agroeca brunnea 27/03/1993 003736/003 em
CLUBIONIDAE (Mouse Spiders)
Comm Clubiona corticalis 29/06/1991 003374/002 em
Comm Clubiona reclusa 06/06/1993 003841/004 em
Comm Clubiona terrestris 01/06/1986 000664/001 em
Comm Clubiona lutescens 06/06/1993 003833/001 em
Comm Clubiona comta 30/05/2009 016756/005 em
Comm Clubiona brevipes 13/07/1992 003617/005 em
GNAPHOSIDAE (Hunter Spiders)
S/an Scotophaeus blackwalli 06/06/1993 003850/002 em
Loc Zelotes latreillei 12/08/1991 003396/004 em
Loc Zelotes apricorum 04/06/1991 003358/007 em
ZORIDAE
Comm Zora spinimana 13/05/1991 003324/010 em
PHILODROMIDAE (Elongate Crab Spiders)
Comm Philodromus dispar 06/06/1994 004164/005 em
Comm Philodromus aureolus 06/06/1993 003837/001 em
Comm Philodromus cespitum 09/07/1994 004189/001 em
Comm Tibellus oblongus 30/05/2009 016762/013
THOMISIDAE (Crab Spiders)
Comm Xysticus cristatus 01/06/1986 000664/003 em
Comm Ozyptila praticola 31/07/2009 016836/011
Loc Ozyptila simplex 30/06/2009 016791/006 em
SALTICIDAE (Jumping Spiders)
Comm Euophrys frontalis 29/06/1991 003397/004 em
3. List of beetles recorded during this survey
Carabidae Ground Beetles
Amara ovata 30/05/09 em 016754
Bembidion lampros 30/05/09 em 016754
Calathus rotundicollis (=piceus) 31/07/09 em 016835
Demetrias atricapillus 30/05/09 em 016762
Leistus ferrugineus 30/10/09 em 016920
Leistus fulvibarbis 30/10/09 em 016924
Loricera pilicornis 30/05/09 em 016754
Nebria brevicollis 30/10/09 em 016916
Notiophilus biguttatus 30/05/09 em 016754
Notiophilus palustris 30/05/09 em 016762
Notiophilus rufipes 30/05/09 em 016754
Oxytelus laqueatus 30/05/09 em 016762
Nb Platyderus depressus 30/05/09 em 016754
Pterostichus strenuus 30/05/09 em 016754
Trechus obtusus 30/06/09 em 016801
Trechus quadristriatus 30/05/09 em 016754
Hydrophilidae
Megasternum concinnum 30/05/09 em 016762
Staphylinidae Rove Beetles
N(b?) Anotylus mutator 27/09/09 em 016888
Anotylus rugosus 30/05/09 em 016762
Anotylus sculpturatus 30/05/09 em 016762
Atheta aquatica 30/05/09 em 016762
Atheta graminicola 30/06/09 em 016795
Atheta hypnorum 31/07/09 em 016828
Autalia impressa 30/10/09 em 016918
Callicerus rigidicornis 31/07/09 em 016836
Lathrobium brunnipes 30/06/09 em 016795
Nb Liogluta pagana 30/05/09 em 016754
Mocyta (Atheta) fungi 30/05/09 em 016762
Mycetoporus lepidus 30/06/09 em 016795
Mycetoporus longulus 30/06/09 em 016795
Mycetota laticollis 31/07/09 em 016828
Ocypus olens 31/07/09 em 016836
Oligota punctulata 30/10/09 em 016916
Olophrum piceum 30/10/09 em 016916
Omalium caesum 30/10/09 em 016916
Omalium italicum 30/10/09 em 016918
Omalium rivulare 30/05/09 em 016754
Othius angustus (=melanocephalus) 30/10/09 em 016923
Othius laeviusculus 30/05/09 em 016754
Othius punctulatus 30/05/09 em 016754
Oxypoda acuminata (lividipennis) 30/05/09 em 016754
Philonthus laminatus 28/08/09 em 016868
Plataraea brunnea 30/05/09 em 016762
Proteinus brachypterus 30/10/09 em 016918
Quedius curtipennis 28/08/09 em 016868
Quedius levicollis (tristis) 27/09/09 em 016888
Quedius picipes 27/09/09 em 016888
Quedius scintillans 30/05/09 em 016754
Rugilus rufipes 30/05/09 em 016754
Sepedophilus nigripennis 31/07/09 em 016828
Stenus impressus 30/10/09 em 016916
Tachyporus chrysomelinus 30/05/09 em 016754
Tachyporus dispar 30/05/09 em 016754
Tasgius morsitans 28/08/09 em 016868
Xantholinus linearis 30/06/09 em 016795
Scydmaenidae
Cephennium gallicum 31/07/09 em 016828
Latridiidae
Corticarina similata 30/05/09 em 016762
Dienerella separanda 28/08/09 em 016868
Enicmus transversus 31/07/09 em 016829
Cryptophagidae
Cryptophagus setulosus 30/06/09 em 016795
Scarabaeidae Scarab Beetles
Serica brunnea 31/07/09 em 016828
Elateridae Click Beetles
Agriotes pallidulus 30/05/09 em 016754
Nb Athous campyloides 31/07/09 em 016828
Lampyridae
Phosphaenus hemipterus 30/06/09 em 016801
Cantharidae
Malthinus flaveolus 30/06/09 em 016801
Rhagonycha fulva 28/08/09 em 016868
Rhagonycha lignosa 30/05/09 em 016762
Chrysomelidae Flower Beetles
Psylliodes affinis 30/05/09 em 016754
Tenebrionidae
Lagria hirta 30/06/09 em 016795
Leiodidae
Nargus velox 30/05/09 em 016754
Nargus wilkini 30/10/09 em 016916
Curculionidae Weevils
Archarius salicivorus 30/05/09 em 016762
Barypeithes pellucidus 30/05/09 em 016754
Curculio glandum 30/05/09 em 016754
Euophryum confine 28/08/09 em 016853
Hypera postica 30/05/09 em 016754
Phyllobius pyri 30/05/09 em 016754
Strophosoma melanogrammum 30/05/09 em 016754
Rhizophagidae
Rhizophagus dispar 30/05/09 em 016754
4. List of all beetles recorded from Coldfall Wood (by Edward Milner)
Carabidae Ground Beetles
Agonum dorsale em
Amara ovata 30/05/09 em 016754
Bembidion lampros 30/05/09 em 016754
Calathus melanocephalus s s em
Calathus rotundicollis (=piceus) em
Demetrias atricapillus 30/05/09 em 016762
Harpalus rufipes em
Leistus ferrugineus 30/10/09 em 016920
Leistus fulvibarbis 30/10/09 em 016924
Leistus rufomarginatus em
Loricera pilicornis EM
Nebria brevicollis em
Notiophilus biguttatus 30/05/09 em 016754
Notiophilus palustris 30/05/09 em 016762
Notiophilus rufipes em
Oxytelus laqueatus 30/05/09 em 016762
Patrobus atrorufus EM
Nb Platyderus depressus 30/05/09 em 016754
Pterostichus madidus em
Pterostichus strenuus em
Trechus obtusus em
Trechus quadristriatus em
Hydrophilidae
Megasternum concinnum em
Staphylinidae Rove Beetles
N(b?) Anotylus mutator 27/09/09 em 016888
Anotylus rugosus 30/05/09 em 016762
Anotylus sculpturatus 30/05/09 em 016762
Anthobium atrocephalum em
Anthobium unicolor em
Atheta aquatica 30/05/09 em 016762
Atheta brunnea em
Atheta graminicola 30/06/09 em 016795
Atheta hypnorum 31/07/09 em 016828
Autalia impressa 30/10/09 em 016918
Callicerus rigidicornis 31/07/09 em 016836
Lathrobium brunnipes 30/06/09 em 016795
Liogluta pagana em
Mocyta (Atheta) fungi em
Mycetoporus lepidus 30/06/09 em 016795
Mycetoporus longulus 30/06/09 em 016795
Mycetota laticollis 31/07/09 em 016828
Ocypus olens 31/07/09 em 016836
Oligota punctulata 30/10/09 em 016916
Olophrum piceum 30/10/09 em 016916
Omalium caesum 30/10/09 em 016916
Omalium italicum 30/10/09 em 016918
Omalium rivulare 30/05/09 em 016754
Othius angustus (=melanocephalus) 30/10/09 em 016923
Othius laeviusculus 30/05/09 em 016754
Othius punctulatus em
Othius subuliformis em
Oxypoda acuminata (lividipennis) em
Philonthus laminatus 28/08/09 em 016868
Plataraea brunnea 30/05/09 em 016762
Proteinus brachypterus 30/10/09 em 016918
Quedius curtipennis 28/08/09 em 016868
Quedius levicollis (tristis) 27/09/09 em 016888
Quedius picipes em
Quedius schatzmayri em
Quedius scintillans 30/05/09 em 016754
Rugilus rufipes 30/05/09 em 016754
Sepedophilus nigripennis 31/07/09 em 016828
Stenus clavicornis em
Stenus impressus 30/10/09 em 016916
Tachinus marginellus em
Tachyporus chrysomelinus 30/05/09 em 016754
Tachyporus dispar 30/05/09 em 016754
Tasgius morsitans em
Xantholinus linearis EM
Scydmaenidae
Cephennium gallicum 31/07/09 em 016828
Nitidulidae
Brachypterus glaber em
Latridiidae
Aridius nodifer em
Corticarina similata 30/05/09 em 016762
Dienerella separanda 28/08/09 em 016868
Enicmus transversus 31/07/09 em 016829
Cryptophagidae
Cryptophagus punctipennis em
Cryptophagus setulosus 30/06/09 em 016795
Micrambe vini em
Byrrhidae Pill Beetles
Byrrhus pilula em
Simplocaria semistriata em
Scarabaeidae Scarab Beetles
Serica brunnea EM
Elateridae Click Beetles
Agriotes obscurus em
Agriotes pallidulus 30/05/09 em 016754
Nb Athous campyloides 31/07/09 em 016828
Athous haemorrhoidalis em
Lampyridae
Phosphaenus hemipterus 30/06/09 em 016801
Cantharidae
Malthinus flaveolus 30/06/09 em 016801
Rhagonycha fulva 28/08/09 em 016868
Rhagonycha lignosa 30/05/09 em 016762
Chrysomelidae Flower Beetles
Aphthona euphorbiae em
Chaetocnema hortensis em
Psylliodes affinis 30/05/09 em 016754
Tenebrionidae
Lagria hirta 30/06/09 em 016795
Leiodidae
Catops nigricans em
Nargus velox em
Nargus wilkini 30/10/09 em 016916
Curculionidae Weevils
Archarius salicivorus 30/05/09 em 016762
Barypeithes araneiformis em
Barypeithes pellucidus 30/05/09 em 016754
Curculio glandum 30/05/09 em 016754
Euophryum confine 28/08/09 em 016853
Hypera postica 30/05/09 em 016754
Otiorhynchus singularis em
Otiorhynchus sulcatus em
Phyllobius pyri 30/05/09 em 016754
Strophosoma melanogrammum em
Rhizophagidae
Rhizophagus dispar 30/05/09 em 016754
5. List of other invertebrates recorded from Coldfall Wood
(all by EM except where stated)
Hemiptera (true bugs)
Tenthredinidae (sawflies)
Athalia rosae 31/07/2009 016832
Lygaeidae (seed bugs, chinch bugs and ground bugs)
Drymus brunneus 31/07/2009
Drymus ryei (T Bantock)
Diptera (flies)
Limoniidae (craneflies)
Limonia nubeculosa
Trichoceridae (winter craneflies)
Trichocera saltator
Trichocera annulata
Panorpidae (scorpion-flies)
Panorpa germanica
Bibionidae (March flies)
Bibio varipes
Asilidae (robber flies)
Dioctria linearis
Rhagionidae (snipe flies)
Rhagio lineola
Hymenoptera (bees, wasps and ants)
Myrmica ruginodis 1991
Pseudoscorpiones (pseudoscorpions)
Chthonius ischnosceles 03/11/1991 003404
Pselaphochernes dubius 30/06/2009 016795
Roncus lubricus 02/07/1990 003727
Diplopoda (millipedes)
Julus scandinavius 22/10/1994 004311
Polydesmus angustus 22/10/1994 004311
Chilopoda (centipedes)
Haplophilus subterraneus 22/10/1994 004311
Lithobius forficatus 22/10/1994 004311
